王寒茹,李丕军,刘华,邢文曦,郑崇文
WANG Han-ru,LI Pi-jun,LIU Hua,XING Wen-xi,ZHENG Chong-wen

• 1:四川省林业科学研究院
• 2:安徽农业大学

摘要(Abstract)：

核桃是世界上著名干果,也是优质硬阔木材树种,主要分布在干旱、半干旱、湿润、半湿润地区,均会受到不同干旱程度的影响。干旱作为一种水分胁迫,对植物的影响主要表现在生长发育及其生理代谢过程。本文系统的综述了国内外关于核桃在干旱条件下核桃形态结构、组织水分状况、质膜系统、渗透调节物质和保护酶系统的变化情况,以期对筛选和培育抗旱高产的核桃品种,对核桃产业的可持续发展提供理论依据。干旱是植物经常遭受的一种逆境,全世界干旱、半干旱地区占全球的43%,在湿润、半湿润地区有时也会出现一定程度的夏旱。干旱对经济林的影响非常大,特别是在干旱胁迫下,对核桃的生长、产量以及坚果质量都有较大影响。为了了解不同核桃品种的抗旱能力,需要了解核桃在干旱条件下生理生化响应,探索机制。筛选和培育抗旱高产的核桃品种,对核桃产业的可持续发展提供理论依据。
The walnut was one of the world's famous dried fruits. also a hardwood species with high quality. It was mainly distributed in arid,semi-arid,humid and semi-humid areas,and affected by different degrees of drought. Drought,as a water stress,impacted on plants mainly in the growth,development and physiological metabolic process. Native and foreign researches were systematically reviewed in the changes of walnut morphology,tissue water status,plasma membrane system,osmotic adjustment substances and protective enzyme system under drought conditions. In order to provide a theoretical basis for screening and breeding of drought-resistant and high-yield varieties of walnuts,and the sustainable development of walnut industry. Drought was an adversity that plants often suffered,the world's arid and semi-arid regions accounted for 43% of the world's total area,and sometimes there would be a certain degree of summer drought in the humid and semi humid areas. The impact of drought on economic forests was very large,especially on the growth,yield and quality of nuts. In order to acquaint the different varieties of walnut drought resistance,it was necessary to understand the physiological and biochemical response of the walnuts in drought conditions,and to explore the mechanism. It could provide a theoretical basis for sustainable development of walnut industry to screen and breed drought-resistant high-yield varieties of the walnut.

关键词(KeyWords)： 干旱;核桃;抗旱性
Drought;Walnut;Drought resistance

Abstract:

Keywords:

基金项目(Foundation): 突破性林木育种材料与方法创新,编号2016NYZ0035;; 核桃耐涝抗旱砧木选择研究,编号2018JBKY07;; 四川核桃主栽品种选择研究,编号2018CZZX17

作者(Author): 王寒茹,李丕军,刘华,邢文曦,郑崇文
WANG Han-ru,LI Pi-jun,LIU Hua,XING Wen-xi,ZHENG Chong-wen

DOI: 10.16779/j.cnki.1003-5508.2018.04.003

参考文献(References)：

• [1]蔡利娟.不同核桃类群综合性状的比较研究[D].四川农业大学,2014.
• [2]陈虹.新疆早实核桃果实生长发育影响因素[D].新疆农业大学,2016.
• [3]孙凡,赵靖明,姚小华,等.水分胁迫逆境处理对水土保持经济树种长山核桃生理特性的影响[J].西南大学学报(自然科学版),2014,36(4):1~7.
• [4]王振元.不同核桃品种抗旱性初步研究[D].西北农林科技大学,2014.
• [5]田艳.干旱胁迫下光核桃根系响应机制初步研究[D].东北林业大学,2016.
• [6]刘杜玲,彭少兵,孙红梅,等.早实核桃不同品种抗旱性综合评价[J].园艺学报,2014,41(5):967~974.
• [7]孙红梅.早实核桃对低温和干旱胁迫的生理响应及抗逆性综合评价[D].西北农林科技大学,2012.
• [8]白重炎,高尚风,张颖,等.12个核桃品种叶片解剖结构及其抗旱性研究[J].西北农业学报,2010,19(7):125~128.
• [9]孙志超,程慧,郭文磊,等.山核桃枝叶耐旱特性分析[J].果树报,2015,32(04):633~640+738.
• [10]王海珍,梁宗锁.黄土高原乡土树种抗旱生理指标的主成分分析[J].塔里木农垦大学学报,2004,16(1):13~15.
• [11]Marshall J,Rutledge R,Blumwald E,et al.Reduction in turgid water volumein jack pine,white spruce and black spruce in response to drought and paclobutrazol.Tree Physiology,2000,20:701~707.
• [12]曹铁森,许明宪,王维友,等.水分胁迫对梨幼树叶水势的影响[J].西北农林科技大学学报(自然科学版),1992,20(1):91~94.
• [13]陈永坤,汪宇.PEG模拟干旱胁迫对漾濞核桃幼苗抗性物质的影响[J].西南林业大学学报,2013,33(4):103~106.
• [14]韩瑞宏,田华,高桂娟,等.干旱胁迫下紫花苜蓿叶片水分代谢与两种渗透调节物质的变化[J].华北农学报,2008,23(4):140~144.
• [15]何海洋,彭方仁,张瑞,等.不同品种美国山核桃嫁接苗光合特性比较[J].南京林业大学学报(自然科学版),2015,39(04):19~25.
• [16]曹丹,陈道钳,吴茜,等.复水对旱后不同玉米品种植株生长恢复能力及其生理响应特性的影响[J].西北植物学报,2015,35(6):1222~1228.
• [17]Khoyerdi FF,Shamshiri MH,Estaji A,et al.Changes in some physiological and osmotic parameters of several pistachio genotypes under drought stress.SCIENTIA HORTICULTURAE,2015,198(10):44~51.
• [18]刘丽君,王以芝,尹田夫.根际干旱对大豆幼苗细胞质膜相对透性及生物产量的影响[J].大豆科学,1986,5(2):117~122.
• [19]胡学华.核桃抗旱生理研究[J].内江科技,2006,1(9):91.
• [20]余甜,徐敏,张萍,等.新疆野核桃抗旱性研究[J].江苏农业科学,2016,44(1):194~196.
• [21]史宝胜,刘冬云,张晓磊,等.水分胁迫对金叶榆含水量、细胞质膜相对透性和抗氧化系统的影响[J].西北农业学报,2010,19(9):88~92.
• [22]Rao N,Sarma P,Chander S.real-time adaptive irrigation scheduling under a limited water suppli.Agricultural water Management,1992,20(4):267~279.
• [23]侯栋,马风云,王笛.干旱胁迫对4核桃品种生化指标的影响[J].西南林学院学报,2010,30(3):24~27.
• [24]李合生.现代植物生理学[M],北京:高等教育出版社,2006.
• [25]Naser,L.V.Kouroshk.Bahman A.Reza.Soluble sugars and proline accumulation paly a role as effective indices for drought tolerance screening in persian walnut(Juglans regia L.)duringgermination.Fruits,2010,65(2):97~112.
• [26]刘济明,赵晓鹏,廖小峰,等.罗甸小米核桃抗氧化保护酶活性和丙二醛含量对干旱胁迫的响应[J].河南农业科学,2012,41(9):122~126.
• [27]A.Rosati,S.Metcalf,R.Buchner,A.Fulton&B.Lampinen.Tree water status and gas exchange in walnut under drought,high temperature and vapour pressure deficit.July,2005,81(3):415~420.
• [28]Lopes MS,Araus JL,Heerden PD,et al.Enhancing drought tolerance in C(4)crops[J].Journal Of Experimental Botany,2011,62(9),3135~3153.
• [29]Golldack D,Luking L,Yang O.Plant tolerance to drought and salinity:stress regulating transcription Factors and their functional significance in the cellular transcriptional network[J].Plant Cell Reports,2011,30(8),1383~1391.
• [30]李长江,崔晓燕,孙帆,等.青杄干旱诱导基因Pw WDS1的c DNA分离与表达分析[J].林业科学,2014,50(04):129~136.
• [31]朱双.麻疯树干旱胁迫基因表达谱分析[D].华南植物园,2012.
• [32]Di Wang,Yajiao Pan,Xiuqin Zhao.Genome-wide temporal-spatial gene expression profiling of drought responsiveness in rice[J].BMC Genomic,2011,12:149.
• [33]Daniela Jerszurki,Valentin Couvreur,Toby Maxwell,Lucas de Carvalho Ramos Silva,Nicholas Matsumoto,Kenneth Shackel,Jorge Luiz Moretti de Souza,Jan Hopmans.Impact of root growth and hydraulic conductance on canopy carbon-water relations of young walnut trees(Juglans regia L.)under drought[J].Scientia Horticulturae,2017.
• [34]K Vahdati,N Lotfi.Abiotic Stress Tolerance in Plants with Emphasizing on Drought and Salinity Stresses in Walnut[M].Abiotic Stress-Plant Responses and Applications in Agriculture,2013.